diff --git a/pyproject.toml b/pyproject.toml
index 93116bede8..88542b2cf5 100644
--- a/pyproject.toml
+++ b/pyproject.toml
@@ -32,7 +32,6 @@ dependencies = [
     'zstandard',
     'typing_extensions',
     'donfig',
-    'pytest'
 ]
 dynamic = [
   "version",
@@ -55,6 +54,22 @@ license = {text = "MIT License"}
 keywords = ["Python", "compressed", "ndimensional-arrays", "zarr"]
 
 [project.optional-dependencies]
+test = [
+    "coverage",
+    "pytest",
+    "pytest-cov",
+    "msgpack",
+    "lmdb",
+    "s3fs",
+    "pytest-asyncio",
+    "moto[s3]",
+    "flask-cors",
+    "flask",
+    "requests",
+    "mypy",
+    "hypothesis"
+]
+
 jupyter = [
     'notebook',
     'ipytree>=0.2.2',
@@ -108,22 +123,7 @@ dependencies = [
     "numpy~={matrix:numpy}",
     "universal_pathlib"
 ]
-extra-dependencies = [
-    "coverage",
-    "pytest",
-    "pytest-cov",
-    "msgpack",
-    "lmdb",
-    "s3fs",
-    "pytest-asyncio",
-    "moto[s3]",
-    "flask-cors",
-    "flask",
-    "requests",
-    "mypy",
-    "hypothesis"
-]
-features = ["extra"]
+features = ["test", "extra"]
 
 [[tool.hatch.envs.test.matrix]]
 python = ["3.10", "3.11", "3.12"]